This is a question that has come up a few times now, and there is a really quick answer : they use two, Lima-style D-ring couplings that are moulded onto the rolling stock (loco, wagon, carriage, etc.) and NEM pocket couplings with reasonably standard tension lock couplings inserted.
The Hornby Railroad (cheaper) range have the D-ring couplings, and the Hornby non-Railroad ranges have the NEM couplings. If in doubt, simply use the product code to search the Hornby shop. If the product has NEM couplings, it will say so under 'Special Features'.
However, there are a few points that I suspect lurk behind the original question. The first related to compatibility.
The old Lima style D-rings are moulded onto the rolling stock, so that they are part of the chassis. This means that changing them to NEM couplings, whilst doable, is a messy process.
You'd be better off buying the more expensive, and slightly more attractive (neither are actually prototypical) models with NEM couplings if that's your goal. Of course, there's a caveat.
That caveat is that Hornby actually sell packs of NEM compatible D-ring couplings. These enable you to buy up Railroad (cheaper) wagons and carriages and then couple them to Hornby non-Railroad locomotives, by taking out the tension lock couplings and replacing them with the D-ring ones.
That's part of the beauty of NEM couplings - you can change the actual couplings for others. This becomes necessary, even if you've splashed out on non-Railroad stock, if you use Brian Kirby style magnetic uncoupling.
The reason is that the Hornby tension lock couplings are made of ferrous material, unlike the Bachmann ones (for example) which are not. If you have ferrous tension lock couplings, then the Brian Kirby approach doesn't work, as the magnets affect the coupling directly.
But that's the beauty of the NEM couplings - if you really don't like them, you can always swap them for some Kadee knuckle couplings, which are generally thought of as being more prototypical, and provide excellent delayed action magnetic uncoupling/coupling which is very versatile.
So, that's pretty much everything I now about the current coupling system that Hornby uses in both the high-end and Railroad product ranges.
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